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Fig. 1. Papillae, beads and axial piles of lamellae. A: papilla and pores. Stereograph, schematic, not to scale; after Hottinger, 1977. B-C: Assilina madagascariensis (d'Orbigny) from Mauritius. Recent. Transverse section parallel to equatorial plane (B) and axial section (C) with minute papillae on the last whorl; transmitted light micrograph. D-F: Beading in Amphistegina papillosa Said. Gulf of Aqaba, Red Sea; Recent. D: dorsal view of shell showing beading of septular and hemiseptular sutures. Incident light micrograph. E: axial section showing hemiseptular and septular support of beads; transmitted light micrograph. F: Epoxy resin cast of shell cavity showing spiral main chamber lumina in dorsal view, interrupted by the hemiseptula supporting the interseptal beads, SEM graph. a: aperture; bd: beads (septular and hemiseptular); ch: main (spiral) chamber lumen; egh: eggholders; hs: hemiseptulum; il: inner lamella; isc: intraseptal canal system; ml: median layer; ol: outer lamellae; orgl: organic lining of protoplast; p: pores; pap: papillae; pil: axial pile of lamellae; s: septum and septal suture; wsut: whorl suture; (Hottinger, 2006; fig. 73 [1] CC/BY-NC-SA)


  • according to Hottinger (2006):

BEAD - a small, rounded to hemispherical protuberance on the surface of lamellar shells, forming strings along septal, septular or hemiseptular sutures. Usually imperforate or poorly perforate. To be distinguished from papilla.

See also


Hottinger (1977), Foraminifères operculiniformes. Mémoires du Muséum National d'Histoire Naturelle, Paris, (Série C, Sciences de la terre), t. XL, 159 p. + 66 pls.

Hottinger (2006), Illustrated glossary of terms used in foraminiferal research. Carnets de Géologie, Memoir 2, ISSN 1634-0744

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